Pachychilidae
Family of gastropods
title: "Pachychilidae" type: doc version: 1 created: 2026-02-28 author: "Wikipedia contributors" status: active scope: public tags: ["pachychilidae", "gastropod-families"] description: "Family of gastropods" topic_path: "general/pachychilidae" source: "https://en.wikipedia.org/wiki/Pachychilidae" license: "CC BY-SA 4.0" wikipedia_page_id: 0 wikipedia_revision_id: 0
::summary Family of gastropods ::
| fossil_range = Paleogene - recent | image = Pachychilus laevisimus 004.JPG | image_caption = Two shells of Pachychilus laevissimus | taxon = Pachychilidae | authority = P. Fischer & Crosse, 1892 | diversity_link = | diversity_ref = | diversity = 191-226 extant species | type_genus = Pachychilus | type_genus_authority = I. Lea & H. C. Lea, 1851 | synonyms_ref = | synonyms = * Brotiinae Golikov & Starobogatov, 1987
- Fauninae Cossmann, 1909
- Melanatriinae Thiele, 1921
- Potadomatinae Pilsbry & Bequaert, 1927 | display_parents = 3
Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.
Distribution
::figure[src="https://upload.wikimedia.org/wikipedia/commons/5/51/Pachychilidae_world.jpg" caption="The global distribution of the family Pachychilidae"] ::
Pachychilids are freshwater snails with a worldwide distribution in the tropics. Representatives are found in South and Central America, Africa, Madagascar, South and South-east Asia and tropical Australia (Queensland: Torres Strait Islands).
Description
Pachychilids have an operculum, which is concentric and multispiral.
Ecology
All species in the family inhabit freshwater except Faunus ater, which is a brackish water snail found in estuaries and other coastal habitats. Pachychilids are either oviparous (lay eggs), ovoviviparous or viviparous (retain developing eggs and youngs in special incubatory structures).
Notes on the taxonomy
The name is derived from a combination of the words 'pachy' (Greek = thick) and the suffix '-chilus' (Greek = Lip), meaning 'thick lipped' - with respect to the thickened aperture of the shell in some species. Most 20th-century authors did not recognize Pachychilidae as an independent family, but affiliated species under different groups, such as Thiaridae and Pleuroceridae. However, recent revisions based on molecular and morphological evidence have proven their independent and distinct status from the former. This family has no subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).
Genera
::figure[src="https://upload.wikimedia.org/wikipedia/commons/5/5b/Pachychilidae_shells.jpg" caption="Shells of various pachychilid species"] ::
Genera within the family Pachychilidae include:
- † Bellatara Strand, 1928 - with only one species Bellatara janus (Mayer, 1870)
- Brotia H. Adams, 1866
- Doryssa Swainson, 1840
- † Eginea Pacaud & Harzhauser, 2012
- Faunus de Montfort, 1810 with the only species Faunus ater (Linnaeus, 1758)
- Jagora Köhler & Glaubrecht, 2003
- † Jponsia Pacaud & Harzhauser, 2012
- Madagasikara Köhler & Glaubrecht, 2010 - synonym: Melanatria Bowdich, 1822 auctt.
- † Nodifaunus Olsson, 1944
- Pachychilus I. Lea & H. C. Lea, 1851 - type genus of the family Pachychilidae
- Paracrostoma Cossmann, 1900
- Potadoma Swainson, 1840
- † Pseudobellardia Cox, 1931
- Pseudopotamis Martens, 1894
- Sulcospira Troschel, 1858 - synonym: Adamietta Brandt, 1974
- † Tinnyea Hantken, 1887
- Tylomelania Sarasin & Sarasin, 1897
- † Wingeastonia K. Martin, 1906 ;Genera brought into synonymy:
- Acrostoma Brot, 1871: synonym of Paracrostoma Cossmann, 1900
- Antimelania P. Fischer & Crosse, 1892: synonym of Brotia H. Adams, 1866
- Brotella Rovereto, 1899: synonym of Paracrostoma Cossmann, 1900
- Fauninae Cossmann, 1909 (subfamily) : synonym of Pachychilidae P. Fischer & Crosse, 1892
- Melanatria Bowdich, 1822: synonym of Faunus Montfort, 1810
- Oxymelania Crosse & P. Fischer, 1892: synonym of Pachychilus I. Lea & H. C. Lea, 1851
- Pirena Lamarck, 1822: synonym of Faunus Montfort, 1810
- Sphaeromelania Rovereto, 1899: synonym of Pachychilus I. Lea & H. C. Lea, 1851
- Wanga Chen, 1943: synonym of Brotia H. Adams, 1866
Cladogram
A cladogram based on sequences of mitochondrial 16S ribosomal RNA sequences showing phylogenic relations of Pachychilidae:
|label1=Pachychilidae |1={{clade |1=Faunus ater |2={{clade |1={{clade |1=Potadoma freethi |2=Doryssa consolidata |2={{clade |1={{clade |1=Sulcospira hainanensis |2={{clade |1=Sulcospira testudinaria |2=Sulcospira housei |3={{clade |label1=Brotia |1={{clade |1=Brotia pagodula |2=Brotia sumatrensis |label2=Paracrostoma |2={{clade |1=Paracrostoma martini |2=Paracrostoma tigrina |2={{clade |1={{clade |1={{clade |label1=Pseudopotamis |1={{clade |1=Pseudopotamis supralirata |2=Pseudopotamis semoni |label2=Tylomelania |2={{clade |1=Tylomelania perfecta |2=Tylomelania neritiformis |label2=Jagora |2={{clade |1=Jagora dactylus |2=Jagora asperata |2={{clade |label1=Pachychilus |1={{clade |1=Pachychilus indiorum |2=Pachychilus conicus |label2=Madagasikara |2={{clade |1=Madagasikara madagascariensis |2={{clade |1=Madagasikara spinosa |2={{clade |1=Madagasikara zazavavindrano |2={{clade |1=Madagasikara vazimba |2=Madagasikara vivipara
References
References
- [[Paul Henri Fischer. Fischer P.]] & [[Joseph Charles Hippolyte Crosse. Crosse H.]] (19 November 1892). "Mission scientifique au Mexique et dans l'Amérique Centrale". ''Recherches zoologiques'', Partie 7, '''2'''(13): 313.
- Strong E. E., Gargominy O., [[Winston Ponder. Ponder W. F.]] & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". ''[[Hydrobiologia]]'' 595: 149-166. http://hdl.handle.net/10088/7390 {{doi. 10.1007/s10750-007-9012-6.
- Strong E. E., Colgan D. J., Healy J. M., Lydeard C., Ponder W. F. & Glaubrecht M. (2011). "Phylogeny of the gastropod superfamily Cerithioidea using morphology and molecules". ''[[Zoological Journal of the Linnean Society]]'' '''162'''(1): 43-89. {{doi. 10.1111/j.1096-3642.2010.00670.x.
- Golikov & Starobogatov (1987). ''Vsesoiuznoe soveshchanie po izucheniiu molliuskov'' '''8''': 25.
- Cossmann (1909). ''Essais de paléoconchologie comparrée'' '''8''': 156.
- Thiele (1921). ''Archiv für molluskenkunde'' '''53'''(3): 142.
- [[Henry Augustus Pilsbry. Pilsbry H. A.]] & [[Joseph Charles Bequaert. Bequaert J.]] (1927). "The Aquatic Mollusks of the Belgian Congo. With a geographical and ecological account of Congo malacology". ''[[Bulletin of the American Museum of Natural History]]'' '''53'''(2): 69-602. pages 248 and 272. [http://hdl.handle.net/2246/975 PDF].
- Gofas, S. (2014). Pachychilidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=411651 on 2014-11-20
- {{Bouchet 2005
- Gofas, S. (2015). Pachychilidae P. Fischer & Crosse, 1892. In: MolluscaBase (2015). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=411651 on 2015-10-09
- Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioidea: Pachychilidae)." ''[[Malacologia]]'' '''48''': 159-251.
- Lok A. F. S. L., Ang W. F., Ng P. X., Ng B. Y. Q. & Tan S. K. (2011). "Status and distribution of ''Faunus ater'' (Linnaeus, 1758) (Mollusca: Cerithioidea) in Singapore". ''[[Nature in Singapore]]'' '''4''': 115-121. [http://rmbr.nus.edu.sg/nis/bulletin2011/2011nis115-121.pdf PDF].
- Köhler F. & Glaubrecht M. (2003). "Morphology, reproductive biology and molecular genetics of ovoviviparous freshwater gastropods (Cerithioidea: Pachychilidae) from the Philippines, with description of the new genus ''Jagora''". ''[[Zoologica Scripta]]'' '''32'''(1): 35-59. {{doi. 10.1046/j.1463-6409.2003.00100.x.
- Köhler F. & Glaubrecht M. (2010). "Uncovering an overlooked radiation: molecular phylogeny and biogeography of Madagascar’s endemic river snails (Caenogastropoda: Pachychilidae: ''Madagasikara'' gen. nov.). ''[[Biological Journal of the Linnean Society]]'' '''99''': 867-894. {{doi. 10.1111/j.1095-8312.2009.01390.x.
- Köhler F. & Glaubrecht M. (2007). "Out of Asia and into India - On the molecular phylogeny and biogeography of the endemic freshwater gastropod ''Paracrostoma'' Cossmann, 1900 (Caenogastropoda: Pachychilidae)." ''[[Biological Journal of the Linnean Society]]'' '''91''': 627-651. {{doi. 10.1111/j.1095-8312.2007.00866.x.
- Glaubrecht M. & von Rintelen T. (2003). "Systematics, molecular genetics and historical zoogeography of the viviparous freshwater gastropod ''Pseudopotamis'' (Cerithioidea, Pachychilidae): a relic on the Torres Strait Islands, Australia". ''[[Zoologica Scripta]]'' '''32'''(5): 415-435. {{doi. 10.1046/j.1463-6409.2003.00127.x.
- Köhler F. & Dames C. (2009). "Phylogeny and systematics of the Pachychilidae of mainland Southeast Asia – novel insights from morphology and mitochondrial DNA (Mollusca, Caenogastropoda, Cerithioidea)". ''[[Zoological Journal of the Linnean Society]]'' '''157''': 679-699. {{doi. 10.1111/j.1096-3642.2009.00541.x.
- von Rintelen T. & Glaubrecht M. (2005). "Anatomy of an adaptive radiation: a unique reproductive strategy in the endemic freshwater gastropod ''Tylomelania'' (Cerithioidea: Pachychilidae) on Sulawesi, Indonesia and its biogeographical implications." ''[[Biological Journal of the Linnean Society]]'' '''85''': 513–542. {{doi. 10.1111/j.1095-8312.2005.00515.x.
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