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Dryandra subg. Hemiclidia

Obsolete subgenus within the former genus ''Dryandra''

Dryandra subg. Hemiclidia is an obsolete plant taxon that encompassed material that is now included in Banksia. Published at genus rank as Hemiclidia by Robert Brown in 1830, it was set aside by George Bentham in 1870, but reinstated at subgenus rank by Alex George in 1996. In 2007, all Dryandra species were transferred into Banksia at series rank, and the infrageneric Dryandra taxa, including D. subg. Hemiclidia, were set aside.

According to Brown

In 1810, Robert Brown published the genus Dryandra in his On the Proteaceae of Jussieu. Thirteen species were published, including Dryandra falcata (now Banksia falcata), but no infrageneric arrangement was proffered. Twenty years later, Brown published a further eleven species and the first infrageneric arrangement in his Supplementum primum prodromi florae Novae Hollandiae. By this time, Brown had observed the tendency in D. falcata for one of the two ovules in each follicle to abort, thereafter developing into a winglike appendage to the seed separator. On this basis, he transferred the species into a monotypic genus, which he named Hemiclidia, from the Greek hemi ("half") and perhaps kleidos ("barrier", "means of closing").

Hemiclidia was retained in the 1856 arrangement of Carl Meissner, but discarded by George Bentham in his 1870 revision of the genus. Bentham correctly observed that the abortion of seeds occurs in other Dryandra species, and is a diagnostically unimportant character:

According to George

In 1996, Alex George published a thorough revision of Dryandra in preparation of his monograph on that genus for the Flora of Australia series. He segregated D. falcata and the newly published D. glauca (now Banksia glaucifolia) into a subgenus based on their unusual follicles, which are small, hairy and pliable. Needing a name for the subgenus, George noted that Hemiclidia was available, and had an appropriate type species in D. falcata. He therefore demoted Hemiclidia to subgenus rank as Dryandra subg. Hemiclidia (R.Br.) A.S.George. Its placement and circumscription in George's taxonomic arrangement of Dryandra may be summarised as follows: :'*Genus Dryandra''' (now Banksia ser. *Dryandra'')

::'*Subgenus *Hemiclidia''''' :::*D. falcata* (now *Banksia falcata*) :::*D. glauca* (now *Banksia glaucifolia*) ::Subgenus *Diplophragma* (1 species) This arrangement remained current until 2007, when Austin Mast and Kevin Thiele transferred *Dryandra* into *Banksia*. They were not yet ready, however, to tender an infrageneric arrangement encompassing *Dryandra*, so as an interim measure they transferred *Dryandra* into *Banksia* at series rank. This minimised the nomenclatural disruption of the transfer, but also caused George's rich infrageneric arrangement to be set aside. Thus under the interim arrangements implemented by Mast and Thiele, *D.* subg. *Hemiclidia* has been set aside. ## References ## References 1. Brown, Robert. (1810). ["On the Proteaceae of Jussieu"](https://zenodo.org/record/1447522). *Transactions of the Linnean Society of London*. 2. Brown, Robert. (1830). "Supplementum Primum Prodromi Florae Novae Hollandiae". *Taylor*. 3. (2006). "The Dryandras". *Melbourne: Australian Plants Society (SGAP Victoria); Perth: Wildflower Society of Western Australia*. 4. Meissner, Carl. (1856). "XLV. Hemiclidia". *Sumptibus Sociorum Treuttel et Wurtz*. 5. Bentham, George. (1870). "Dryandra". *L. Reeve & Co.*. 6. George, Alex S.. (1996). ["New taxa and a new infragenetic classification in Dryandra R.Br. (Proteaceae: Grevilleoideae)"](https://www.biodiversitylibrary.org/part/235162). *[[Nuytsia (journal)*. 7. George, Alex S.. (1999). "Banksia". *CSIRO Publishing / Australian Biological Resources Study*. 8. (2007). "The transfer of Dryandra R.Br. to Banksia L.f. (Proteaceae)". *Australian Systematic Botany*. ::callout[type=info title="Wikipedia Source"] This article was imported from [Wikipedia](https://en.wikipedia.org/wiki/Dryandra_subg._Hemiclidia) and is available under the [Creative Commons Attribution-ShareAlike 4.0 License](https://creativecommons.org/licenses/by-sa/4.0/). Content has been adapted to SurfDoc format. Original contributors can be found on the [article history page](https://en.wikipedia.org/wiki/Dryandra_subg._Hemiclidia?action=history). ::

banksia-ser.-dryandra historically-recognized-angiosperm-taxa plant-subgenera taxa-named-by-robert-brown-(botanist,-born-1773)

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