Pakicetus

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title: "Pakicetus" type: doc version: 1 created: 2026-02-28 author: "Wikipedia contributors" status: active scope: public tags: ["transitional-fossils", "pakicetidae", "fossils-of-pakistan", "eocene-mammals-of-asia", "prehistoric-cetacean-genera", "fossil-taxa-described-in-1981", "taxa-named-by-philip-d.-gingerich"] description: "Genus of ancient whales" topic_path: "geography/pakistan" source: "https://en.wikipedia.org/wiki/Pakicetus" license: "CC BY-SA 4.0" wikipedia_page_id: 0 wikipedia_revision_id: 0

::summary Genus of ancient whales ::

| fossil_range = Eocene (Ypresian), | image = Pakicetus Canada.jpg | image_caption = Cast of P. attocki, Canadian Museum of Nature | taxon = Pakicetus | authority = | type_species = †Pakicetus inachus | type_species_authority = | subdivision_ranks = Species | subdivision = * †P. inachus

• †P. attocki

• †P. calcis

• †P. chittas

Pakicetus (meaning 'whale from Pakistan') is an extinct genus of amphibious cetacean of the family Pakicetidae, which was endemic to South Asia during the Ypresian (early Eocene) period, about 50 million years ago. It was a wolf-like mammal, about 1-2 m long,

History of discovery

The first fossil, a skull fragment of P. inachus, was found in 1981 in Pakistan. Subsequent fossils of Pakicetus were also found in Pakistan, hence the generic name Pakicetus. The fossils were found in the Kuldana Formation west of Islamabad in northern Pakistan and were dated as early to early-middle Eocene in age. The discovery of Pakicetus played an important role in solidifying the inferences that revolved around the evolution of whales. The fossil indicated that whales swam up and down with their vertebral column, which caused their feet to move up and down like otters and their land movements were similar to sea lions; even their limbs protracted and retracted on land. In contrast, the origin of cetaceans, which includes whales, began as four-legged land animals who actively used locomotion and were great runners as a result.

The fossils came out of red terrigenous sediments bounded largely by shallow marine deposits typical of coastal environments caused by the Tethys Ocean. Speculation is that many major marine banks flourished with the presence of this prehistoric whale. According to the location of fossil findings, the animals preferred a shallow habitat that neighbored decent-sized land. Assortments of limestone, dolomite, stone mud and other varieties of different coloured sands have been predicted to be a favourable habitat for Pakicetus. During the Eocene, modern day Pakistan was part of an independent island-continent off the coastal region of Eurasia, and therefore an ideal habitat for the evolution and diversification of the Pakicetidae.

Classification

Pakicetus was classified as an early cetacean due to characteristic features of the inner ear found only in cetaceans (namely, the large auditory bulla is formed from the ectotympanic bone only). It was recognized as the earliest member of the family Pakicetidae. Thus, Pakicetus represents a transitional taxon between extinct land mammals and modern cetaceans.

believed Pakicetus to be a mesonychid. However, studies from molecular biology placed today's cetaceans within the group of artiodactyls, to which the mesonychids do not belong. In 2001, fossils of ancient whales were found that featured an ankle bone, the astragalus, with a "double pulley" shape characteristic of artiodactyls. The redescription of the semi-aquatic, small deer-like artiodactyl Indohyus, and the discovery of its cetacean-like inner ear, simultaneously put an end to the idea that whales were descended from mesonychids, while demonstrating that Pakicetus, and all other cetaceans, are artiodactyls.

Description

Based on the sizes of specimens, and to a lesser extent on composite skeletons, species of Pakicetus are thought to have been 1 m to 2 m in length.

Pakicetus looked very different from modern cetaceans, and its body shape more resembled those of land-dwelling hoofed mammals. Unlike all later cetaceans, it had four fully functional long legs. Pakicetus had a long snout; a typical complement of teeth that included incisors, canines, premolars, and molars; a distinct and flexible neck; and a very long and robust tail. As in most land mammals, the nose was at the tip of the snout.

Reconstructions of pakicetids that followed the discovery of composite skeletons often depicted them with fur; however, given their relatively close relationships with hippos, they may have had sparse body hair.

The first fossil found consisted of an incomplete skull with a skull cap and a broken mandible with some teeth. Based on the detail of the teeth, the molars suggest that the animal could rip and tear flesh. Wear, in the form of scrapes on the molars, indicated that Pakicetus ground its teeth as it chewed its food. Because of the tooth wear, Pakicetus is thought to have eaten fish and other small animals. The teeth also suggest that Pakicetus had herbivorous and omnivorous ancestors.

Palaeobiology

::figure[src="https://upload.wikimedia.org/wikipedia/commons/3/34/Pakicetus_BW.jpg" caption="''P. inachus'' life restoration"] ::

Possible semi-aquatic nature

It was illustrated on the cover of Science in 1983 as a semiaquatic, vaguely crocodile-like mammal, diving after fish.

Somewhat more complete skeletal remains were discovered in 2001, prompting the view that Pakicetus was primarily a land animal about the size of a wolf. wrote that "Pakicetids were terrestrial mammals, no more amphibious than a tapir."

However, argued that "the orbits ... of these cetaceans were located close together on top of the skull, as is common in aquatic animals that live in water but look at emerged objects. Just like Indohyus, limb bones of pakicetids are osteosclerotic, also suggestive of aquatic habitat" (since heavy bones provide ballast). "This peculiarity could indicate that Pakicetus could stand in water, almost totally immersed, without losing visual contact with the air."

The dental microwear of P. inachus was similar to that of semiaquatic protocetids from the Middle Eocene, showing that this species was already deriving its dietary needs from aquatic prey items.

Sensory capabilities

The Pakicetus skeleton reveals several details regarding the creature's unique senses and provides a newfound ancestral link between terrestrial and aquatic animals. As previously mentioned, the *Pakicetus''' upward-facing eye placement was a significant indication of its habitat. Even more so, however, was its auditory abilities. Like all other cetaceans, Pakicetus had a thickened skull bone known as the auditory bulla, which was specialized for underwater hearing.

Notes

References

• {{Cite web | last = Polly | first = Paul D. | title = Pakicetus (fossil Mammal Genus) | publisher = Encyclopædia Britannica Online | url = http://global.britannica.com/EBchecked/topic/1700783/Pakicetus | access-date = 24 October 2013 | archive-date = 29 October 2013 | archive-url = https://web.archive.org/web/20131029201255/http://global.britannica.com/EBchecked/topic/1700783/Pakicetus | url-status = live
• {{Cite journal | last1 = Cooper | first1 = Lisa Noelle | last2 = Thewissen | first2 = J. G. M. | author-link2 = Hans Thewissen | last3 = Hussain | first3 = S. T. | title = New middle Eocene archaeocetes (Cetacea: Mammalia) from the Kuldana Formation of northern Pakistan | year = 2009 | journal = Journal of Vertebrate Paleontology | volume = 29 | issue = 4 | pages = 1289–1299 | doi = 10.1671/039.029.0423| bibcode = 2009JVPal..29.1289C | s2cid = 84127292
• {{Cite journal | last = de Muizon | first = Christian | title = L'origine et l'histoire évolutive des Cétacés |trans-title=Origin and evolutionary history of cetaceans |language=fr |date=March 2009 | journal = Comptes Rendus Palevol | volume = 8 | issue = 2–3 | pages = 295–309 | doi = 10.1016/j.crpv.2008.07.002| bibcode = 2009CRPal...8..295D
• {{Cite journal | last1 = Gingerich | first1 = Philip D. | author1-link = Philip D. Gingerich | last2 = Wells | first2 = N. A. | last3 = Russell | first3 = Donald E. | last4 = Shah | first4 = S. M. Ibrahim | title = Origin of Whales in Epicontinental Remnant Seas: New Evidence from the Early Eocene of Pakistan | date = 22 April 1983 | journal = Science | volume = 220 | issue = 4595 | pages = 403–406 | url = http://www-personal.umich.edu/~gingeric/PDFfiles/PDG133_Pakicetus.pdf | access-date = 1 June 2013 | doi = 10.1126/science.220.4595.403 | pmid = 17831411 | bibcode = 1983Sci...220..403G | s2cid = 37602424 }}
• {{Cite journal | last1 = Gingerich | first1 = Philip D. | last2 = Russell | first2 = Donald E. | title = Pakicetus inachus, A New Archaeocete (Mammalia, Cetacea) from the Early-Middle Eocene Kuldana Formation of Kohat (Pakistan) | year = 1981 | journal = Contributions from the Museum of Paleontology, the Museum of Michigan | volume = 25 | issue = 11 | pages = 235–246 | url = http://deepblue.lib.umich.edu/bitstream/handle/2027.42/48501/ID352.pdf | access-date = 1 February 2013 | oclc = 742729300}}
• {{Cite journal | last1 = Gingerich | first1 = Philip D. | title = Stratigraphic and micropaleontological constraints on the middle Eocene age of the mammal-bearing Kuldana Formation of Pakistan | year = 2003b | journal = Journal of Vertebrate Paleontology | volume = 23 | issue = 3 | pages = 643–651 | doi = 10.1671/2409| bibcode = 2003JVPal..23..643G | s2cid = 131407382
• {{Cite journal | last1 = Thewissen | author-link = Hans Thewissen | first1 = J. G. M. | last2 = Williams | first2 = E. M. | last3 = Roe | first3 = L. J. | last4 = Hussain | first4 = S. T. | title = Skeletons of terrestrial cetaceans and the relationship of whales to artiodactyls | year = 2001 | journal = Nature | volume = 413 | issue = 6853 | pages = 277–281 | url = http://www.faculty.virginia.edu/bio202/202-2002/Lectures%2020202/thesissen%20et%20al%202001.pdf | access-date = 1 June 2013 | doi = 10.1038/35095005 | pmid = 11565023| bibcode = 2001Natur.413..277T | s2cid = 4416684
• {{Cite journal | last1 = Thewissen | author-link = Hans Thewissen | first1 = J.G.M. | last2 = Hussain | first2 = S.T. | title = Origin Of Underwater Hearing in Whales | year = 1993 | journal = Nature | volume = 361 | issue = 6411 | pages = 444–445 | doi = 10.1038/361444a0 | pmid = 8429882 | bibcode = 1993Natur.361..444T | s2cid = 4319369
• {{Cite journal | last = West | first = Robert M | title = Middle Eocene large mammal assemblage with Tethyan affinities, Ganda Kas region, Pakistan | year = 1980 | journal = Journal of Paleontology | volume = 54 | issue = 3 | pages = 508–533 | oclc = 4899161959 | jstor = 1304193}}

References

1. (22 December 1998). "A new Eocene archaeocete (Mammalia, Cetacea) from India and the time of origin of whales". Proceedings of the National Academy of Sciences.
2. Black, Riley. (1 December 2010). "How Did Whales Evolve?".
3. (1994). "Fossil Evidence for the Origin of Aquatic Locomotion in Archaeocete Whales". Science.
4. (8 October 2014). "A life spent chasing down how whales evolved".
5. "''Pakicetus'' spp.". NYIT College of Osteopathic Medicine.
6. (2016). "Cetacean Paleobiology". Wiley-Blackwell.
7. {{Harvnb. Gingerich. Wells. Russell. Shah. 1983
8. {{Harvnb. Thewissen. Williams. Roe. Hussain. 2001
9. (15 September 2013). "Paleoecology of archaeocete whales throughout the Eocene: Dietary adaptations revealed by microwear analysis". [[Palaeogeography, Palaeoclimatology, Palaeoecology]].
10. Other cetaceans also all categorically exhibit a large mandibular [[foramen]] within the lower jaw, which holds a fat pack and extends towards the ear, both of which are also associated with underwater hearing. "''Pakicetus'' is the only cetacean in which the mandibular foramen is small, as is the case in all terrestrial animals. It thus lacked the fat pad, and sounds reached its eardrum following the [[external auditory meatus]] as in terrestrial mammals. Thus the hearing mechanism of ''Pakicetus'' is the only known intermediate between that of land mammals and aquatic cetaceans."{{Harvnb. Thewissen. Hussain. 1993

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