Pacullidae

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title: "Pacullidae" type: doc version: 1 created: 2026-02-28 author: "Wikipedia contributors" status: active scope: public tags: ["pacullidae", "araneomorphae-families"] description: "Family of spiders" topic_path: "general/pacullidae" source: "https://en.wikipedia.org/wiki/Pacullidae" license: "CC BY-SA 4.0" wikipedia_page_id: 0 wikipedia_revision_id: 0

::summary Family of spiders ::

| name = | image = | image_caption = | image2 = | image2_caption = | taxon = Pacullidae | authority = Simon, 1894 | subdivision_ranks = | subdivision = | oldest_fossil = Cenomanian | diversity = 4 genera, 38 species | range_map = Pacullidae range map.svg | range_map_caption = blue: reported countries (WSC)

Pacullidae is a family of araneomorph spiders first described by Eugène Simon in 1894. It was merged into Tetrablemmidae in 1958, then raised back to family status after a large phylogenetic study in 2017.

Description

The family Pacullidae contains three-clawed spiders with six eyes, lacking a cribellum. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.

Phylogeny

Pacullidae falls within the Synspermiata clade, a clade of former haplogyne spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to Diguetidae. Together with Pholcidae, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system. |label1="Lost trachea clade" |1={{clade |1={{clade |1=Tetrablemmidae |2={{clade |1=Plectreuridae |2={{clade |1=Diguetidae |2=Pacullidae |2=Pholcidae

Genera

, this family includes four genera and 38 species:

• Lamania Lehtinen, 1981 – Southeast Asia, Bali, Borneo
• Paculla Simon, 1887 – Malaysia, Singapore, New Guinea
• Perania Thorell, 1890 – China, Southeast Asia
• Sabahya Deeleman-Reinhold, 1980 – Borneo

Extinct

• †Furcembolus Wunderlich 2008 9 spp, Burmese amber, Myanmar, Cenomanian

References

References

1. "Family Pacullidae Simon, 1894". World Spider Catalog.
2. Simon, E.. (1894). "Histoire naturelle des araignées".
3. Bonnet, P.. (1958). "Bibliographia araneorum". Douladoure, Toulouse.
4. Brignoli, P. M.. (1973). "Ragni della Melanesia, I. Un nuovo Tetrablemma di Guadalcanal (Isole Salomone) (Araneae Tetrablemmidae)". Memorie della Società Entomologica Italiana, Genova.
5. (2016). "The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling". Cladistics.

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