Gleicheniaceae

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Family of ferns

title: "Gleicheniaceae" type: doc version: 1 created: 2026-02-28 author: "Wikipedia contributors" status: active scope: public tags: ["gleicheniales", "fern-families"] description: "Family of ferns" topic_path: "general/gleicheniales" source: "https://en.wikipedia.org/wiki/Gleicheniaceae" license: "CC BY-SA 4.0" wikipedia_page_id: 0 wikipedia_revision_id: 0

::summary Family of ferns ::

| name = Forked ferns | fossil_range = | image = Diplopterygium pinnatum1.jpg | image_caption = Diplopterygium pinnatum | taxon = Gleicheniaceae | authority = C.Presl | subdivision_ranks = Genera | subdivision =

The forked ferns are the family Gleicheniaceae, which includes eight genera The formerly independent families Dicranopteridaceae and Stromatopteridaceae are generally included in the Gleicheniaceae, whereas the Dipteridaceae and Matoniaceae, although closely related, are considered separate families by most authors.

Description

These tropical ferns are the most widespread living lineage of Gleicheniales. Their rhizomes have a "vitalized" protostele or in some taxa a solenostele. The leaves are indeterminate, with pseudodichotomously forked leaves except in Stromatopteris, and free veins. Stromatopteris also differs from the other species in having an achlorophyllous and subterranean gametophyte. The sori are abaxial but not marginal and carry 5–15 exindusiate round sporangia each. These have a transverse-oblique annulus and contain 128 to 800 bilateral or globose-tetrahedral spores. The sori and sporangia mature at the same time, and the spores grow into surface-dwelling green prothalli beset with club-shaped hairs.

Systematics

There is some dispute about ranking these ferns. They are typically placed in the order Gleicheniales. They may alternatively be considered a subclass Gleicheniatae or class Gleichenopsida, so that a distinct taxon can be established for the leptosporangiate ferns. In this case, the Gleicheniaceae are upranked to an order and receive the name Gleicheniales. Irrespective of their modern taxonomic treatment, the Gleicheniaceae were formerly included in the order Polypodiales. But the ferns in the loose sense are much too diverse a group to be shoehorned into one taxon at such a low rank.

The Triassic Antarctipteris and Gleichenipteris are sometimes ascribed to the Gleicheniaceae, but are probably better considered Gleicheniales incertae sedis.

Phylogeny of Gleicheniaceae by Nitta et al. 2022 and Fern Tree of life |label1=Gleicheniaceae |1={{clade |1=Rouxopteris Liu 2020 |2={{clade |1={{clade |1=Diplopterygium Nakai 1950 |2={{clade |1=Gleichenella Ching 1940 |2=Dicranopteris Bernhardi 1805 |2={{clade |1={{clade |1=Stromatopteris Mettenius 1861 |2=Gleichenia Smith 1973 |2=Sticherus Presl 1836

Footnotes

References

References

  1. "Gleicheniaceae C.Presl". [[Royal Botanic Gardens, Kew]].
  2. (2016). "The number of known plants species in the world and its annual increase". Magnolia Press.
  3. Smith ''et al.'' (2006)
  4. [https://bsapubs.onlinelibrary.wiley.com/doi/10.1002/aps3.11465 Identifying cryptic fern gametophytes using DNA barcoding: A review]
  5. PPG I. (2016). "A community-derived classification for extant lycophytes and ferns". Journal of Systematics and Evolution.
  6. (April 2020). "The significance of Rouxopteris (Gleicheniaceae, Polypodiopsida): a new genus endemic to the Madagascan region". Plant Systematics and Evolution.
  7. (2022). "An Open and Continuously Updated Fern Tree of Life". Frontiers in Plant Science.
  8. (2024). "Tree viewer: interactive visualization of FTOL".

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gleichenialesfern-families