Eupulmonata

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title: "Eupulmonata" type: doc version: 1 created: 2026-02-28 author: "Wikipedia contributors" status: active scope: public tags: ["panpulmonata"] description: "Clade of gastropods" topic_path: "general/panpulmonata" source: "https://en.wikipedia.org/wiki/Eupulmonata" license: "CC BY-SA 4.0" wikipedia_page_id: 0 wikipedia_revision_id: 0

::summary Clade of gastropods ::

| image = Snail-WA.jpg | image_caption = Cepaea sp. | taxon = Eupulmonata | authority = Haszprunar & Huber, 1990 | subdivision_ranks = Orders | subdivision = *Systellommatophora

• Stylommatophora
• Ellobiida

Eupulmonata is a taxonomic clade of air-breathing gastropod molluscs. The great majority of this group are land snails and slugs, but some are intertidal or inhabit coastal saltmarshes and mangroves.

Possible synapomorphy of the group are globineurons (small cells) in the procerebrum.

Eyes positioned on the tips of the tentacles were considered a synapomorphy of a clade named Geophila (Stylommatophora + Systellommatophora), but a 1972 phylogenomic study found strong support for a clade uniting Ellobioidea (eyes at the base of tentacles) and Systelommatopohra (eyes on the tentacle tips). Stalked eyes thus likely evolved twice independently.

Taxonomy

Source:

• Order Ellobiida
  • Superfamily Ellobioidea L. Pfeiffer, 1854
    • Ellobiidae L. Pfeiffer, 1854
    • Otinidae H. Adams & A. Adams, 1855
    • Trimusculidae J. Q. Burch, 1945
• Order Systellommatophora
  • Superfamily Onchidioidea Rafinesque, 1815
    • Onchidiidae Rafinesque, 1815
  • Superfamily Veronicelloidea Gray, 1840
    • Veronicellidae Gray, 1840
    • Rathouisiidae Heude, 1885
• Order Stylommatophora
  • (for inner division, see the Stylommatophora page) ::figure[src="https://upload.wikimedia.org/wikipedia/commons/1/1f/Caucasotachea_atrolabiata.jpg" caption="''[[Caucasotachea atrolabiata]]''"] ::

::figure[src="https://upload.wikimedia.org/wikipedia/commons/d/d3/Helicella_macedonica.jpg" caption="''[[Helicella macedonica]]''"] ::

References

References

1. (2022-04-13). "Phylogenomic resolution of the root of Panpulmonata, a hyperdiverse radiation of gastropods: new insight into the evolution of air breathing". [[Proceedings of the Royal Society B: Biological Sciences]].
2. (2017). "Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families". [[Malacologia]].

::callout[type=info title="Wikipedia Source"] This article was imported from Wikipedia and is available under the Creative Commons Attribution-ShareAlike 4.0 License. Content has been adapted to SurfDoc format. Original contributors can be found on the article history page. ::