Anapidae

Family of spiders
title: "Anapidae" type: doc version: 1 created: 2026-02-28 author: "Wikipedia contributors" status: active scope: public tags: ["anapidae", "araneomorphae-families", "cosmopolitan-spiders", "taxa-named-by-eugène-simon"] description: "Family of spiders" topic_path: "general/anapidae" source: "https://en.wikipedia.org/wiki/Anapidae" license: "CC BY-SA 4.0" wikipedia_page_id: 0 wikipedia_revision_id: 0
::summary Family of spiders ::
| fossil_range = | image = Conoculus.lyugadinus.female.-.tanikawa.jpg | image_caption = C. lyugadinus, female from Okinawa | image2 = Conoculus.lyugadinus.male.-.tanikawa.jpg | image2_caption = C. lyugadinus, male | taxon = Anapidae | authority = Simon, 1895 | diversity = 59 genera, 233 species | synonyms = Micropholcommatidae
Holarchaeidae | range_map = Anapidae range map.svg | range_map_caption = blue: reported countries (WSC)
Anapidae is a family of rather small spiders with 233 described extant species in 59 genera. It includes the former family Micropholcommatidae as the subfamily Micropholcommatinae, and the former family Holarchaeidae. Most species are less than 2 mm long.
They generally live in leaf litter and moss on the floor of rain forest. Many build orb webs with a diameter less than 3 cm. In some species, such as Pseudanapis parocula, the pedipalps of the female are reduced to coxal stumps.
Description
Dippenaaria luxurians side view 01.jpg|Dippenaaria luxurians Dippenaaria luxurians dorsal view.jpg|D. luxurians
Spiders of this family are very small, usually less than two millimeters long, and lack a cribellum. They can have zero, six or eight eyes, the rear median eyes either reduced or missing. One species: 'Epigastrina typhlops' (Rix & Harvey, 2010) has no eyes, an adaption to life underground. In some genera the carapace is modified so that the eyes are raised higher than usual. Color can range from reddish brown, yellowish brown to pale, pigment reduced, creams. Generally both margins of chelicerae have teeth, one exception being 'Acrobleps hygrophilus' who lacks retromarginal teeth. The legs are short and spineless, though there are some with reduced spination such as the genus: 'Teutoniella' or with only one or two long spines such as 'Borneanapis'). The labium has a spur that extends between the chelicerae and can be seen when the chelicerae are spread.
Distribution
Anapidae are found worldwide, particularly in South America, Africa, Asia, Australia and New Zealand. Few genera occur in North America or Europe. Only Comaroma simoni and the three species of Zangherella are found in Europe; Gertschanapis shantzi and Comaroma mendocino are found in the United States.
Systematics
Novanapis spinipes male dorsal.jpg|male Novanapis spinipes Novanapis spinipes male lateral.jpg|male N. spinipes
The family Micropholcommatidae was synonymized with this family by Schütt in 2003 and by Lopa et al. in 2011, a change that has been accepted by the World Spider Catalog. Similarly, the family Holarchaeidae was synonymized by Dimitrov et al. in 2017, and likewise accepted by the World Spider Catalog.
Genera
, this family includes 59 genera and 233 species:
- Acrobleps Hickman, 1979 – Tasmania
- Algidiella Rix & Harvey, 2010 – New Zealand (Auckland Islands)
- Anapis Simon, 1895 – Central America, South America
- Anapisona Gertsch, 1941 – Central America, South America
- Austropholcomma Rix & Harvey, 2010 – Western Australia, Tasmania
- Borneanapis Snazell, 2009 – Borneo
- Caledanapis Platnick & Forster, 1989 – New Caledonia
- Chasmocephalon O. Pickard-Cambridge, 1889 – Australia
- Comaroma Bertkau, 1889 – China, Europe, Japan, Korea, United States
- Conculus Kishida, 1940 – Asia
- Crassanapis Platnick & Forster, 1989 – Argentina, Chile
- Crozetulus Hickman, 1939 – Crozet Islands, Southern Africa
- Dippenaaria Wunderlich, 1995 – South Africa
- Elanapis Platnick & Forster, 1989 – Chile
- Enielkenie Ono, 2007 – Taiwan
- Eperiella Rix & Harvey, 2010 – Tasmania, Chile
- Epigastrina Rix & Harvey, 2010 – Tasmania
- Eterosonycha Butler, 1932 – Australia
- Forsteriola Brignoli, 1981 – Central Africa
- Gaiziapis Miller, Griswold & Yin, 2009 – China
- Gertschanapis Platnick & Forster, 1990 – United States
- Gigiella Rix & Harvey, 2010 – Australia, Chile
- Guiniella Rix & Harvey, 2010 – New Guinea
- Hickmanapis Platnick & Forster, 1989 – Tasmania
- Holarchaea Forster, 1955 – Tasmania, New Zealand
- Mandanapis Platnick & Forster, 1989 – New Caledonia
- Maxanapis Platnick & Forster, 1989 – Australia
- Metanapis Brignoli, 1981 – Kenya, South Africa, Congo, Nepal
- Micropholcomma Crosby & Bishop, 1927 – Australia
- Minanapis Platnick & Forster, 1989 – Argentina, Chile, China
- Montanapis Platnick & Forster, 1989 – New Caledonia
- Normplatnicka Rix & Harvey, 2010 – Australia, Chile
- Nortanapis Platnick & Forster, 1989 – Australia (Queensland)
- Novanapis Platnick & Forster, 1989 – New Zealand
- Octanapis Platnick & Forster, 1989 – Australia
- Olgania Hickman, 1979 – Australia (Tasmania)
- Paranapis Platnick & Forster, 1989 – New Zealand
- Patelliella Rix & Harvey, 2010 – Lord Howe Island
- Pecanapis Platnick & Forster, 1989 – Chile
- Plectochetos Butler, 1932 – Australia
- Pseudanapis Simon, 1905 – Asia, Oceania, Central America, South America. Introduced to Europe, Mauritius, Réunion
- Pua Forster, 1959 – New Zealand
- Queenslanapis Platnick & Forster, 1989 – Australia (Queensland)
- Raveniella Rix & Harvey, 2010 – Australia
- Rayforstia Rix & Harvey, 2010 – Australia, New Zealand
- Risdonius Hickman, 1939 – Australia
- Sheranapis Platnick & Forster, 1989 – Chile
- Sinanapis Wunderlich & Song, 1995 – Southeast Asia
- Sofanapis Platnick & Forster, 1989 – Chile
- Spinanapis Platnick & Forster, 1989 – Australia (Queensland)
- Taliniella Rix & Harvey, 2010 – New Zealand
- Taphiassa Simon, 1880 – Australia, New Caledonia, New Zealand, Sri Lanka
- Tasmanapis Platnick & Forster, 1989 – Tasmania
- Teutoniella Brignoli, 1981 – Brazil, Chile
- Tinytrella Rix & Harvey, 2010 – New Zealand
- Tricellina Forster & Platnick, 1989 – Chile
- Victanapis Platnick & Forster, 1989 – Australia (Victoria)
- Zangherella Caporiacco, 1949 – Southern Europe, Tunisia
- Zealanapis Platnick & Forster, 1989 – New Zealand
There are also genera only known as fossils, typically in Baltic amber: | † Balticoroma | Wunderlich, 2004 | † Balticonopsis | Wunderlich, 2004 | † Cenotextricella | Penney in Penney et al., 2007 | † Deanoorapsis | Penney, 2020 | † Dubianapis | Wunderlich, 2004 | † Flagellanapis | Wunderlich, 2004 | † Fossilanapis | Wunderlich, 2004 | † Palaeoanapis | Wunderlich, 1988 | † Ruganapis | Wunderlich, 2004 | † Saxonanapis | Wunderlich, 2004 | † Tuberanapis | Wunderlich, 2004
References
References
- "Family: Anapidae Simon, 1895". World Spider Catalog.
- "NMBE - World Spider Catalog".
- "Family: Holarchaeidae Forster & Platnick, 1984". Natural History Museum Bern.
- https://britishspiders.org.uk/system/files/library/140907.pdf {{Bare URL PDF. (August 2025)
- (2023). "A summary list of fossil spiders and their relatives, version 23.5". Natural History Museum Bern.
- "Epigastrina typhlops sp. n. from the Mole Creek karst, Tasmania. A, C... | Download Scientific Diagram".
- (2014). "Systematics, Phylogeny, and Evolution of Orb-Weaving Spiders". Annual Review of Entomology.
- (2011). "Morphology to the rescue: molecular data and the signal of morphological characters in combined phylogenetic analyses — a case study from mysmenid spiders (Araneae, Mysmenidae), with comments on the evolution of web architecture". Cladistics.
- (2000). "An introduction to the spiders of South East Asia with notes on all the genera". Malaysian Nature Society Kuala Lumpur.
- (2003). "Phylogeny of Symphytognathidae". Zoologica Scripta.
- (1999). "The Spiders of China". Hebei University of Science and Technology, Publishing House.
- (2008). "Phylogenetic placement of the Tasmanian spider ''Acrobleps hygrophilus'' (Araneae, Anapidae) with comments on the evolution of the capture web in Araneoidea". Cladistics.
- (1986). "On Teutoniella, an American genus of the spider family Micropholcommatidae (Araneae, Palpimanoidea)".
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